Prof Charles Haddad
Associate Professor
Zoology and Entomology
D 208
IB 49

Short CV

PhD (Entomology) - University of the Free State, 2012



Publications (Short List)

Publications (2016-2018) 


CECCARELLI, F.S., HADDAD, C.R. & RAMÍREZ, M.J. 2016. First record of endosymbiontic Rickettsiales (Alphaproteobacteria) from the spider genus Amaurobioides (Araneae: Anyphaenidae). Journal of Arachnology 44: 251–253.


CECCARELLI, F.S., OPELL, B.D., HADDAD, C.R., RAVEN, R.J., SOTO, E.M. & RAMÍREZ, M.J. 2016. Around the World in Eight Million Years: historical biogeography of the intertidal spider genus Amaurobioides (Araneae: Anyphaenidae). PlosONE 11(#e0163740): 1–20.


FOORD, S.H., DIPPENAAR-SCHOEMAN, A.S., JOCQUÉ, R., HADDAD, C.R., LYLE, R. & WEBB, P. 2016. South African National Survey of Arachnida: A checklist of the spiders (Arachnida, Araneae) of the Lekgalameetse Nature Reserve, Limpopo Province, South Africa. Koedoe 58(#a1405): 1–8.

HADDAD, C.R. 2016. Diversity and ecology of spider assemblages associated with Vachellia xanthophloea bark in a South African reserve (Arachnida: Araneae). African Entomology 24: 321–333.

HADDAD, C.R., BRABEC, M., PEKÁR, S. & FOURIE, R. 2016. Seasonal population dynamics of a specialised termite-eating spider (Araneae: Ammoxenidae) and its prey (Isoptera: Hodotermitidae). Pedobiologia 59: 105–110.


JANION-SCHEEPERS, C., MEASEY, J., BRASCHLER, B., CHOWN, S.L., COETZEE, L., COLVILLE, J., DAMES, J., DAVIES, A.B., DAVIES, S., DAVIS, A., DIPPENAAR-SCHOEMAN, A.S., DUFFY, G., FOURIE, D., GRIFFITHS, C., HADDAD, C.R., HAMER, M., HERBERT, D., HUGO-COETZEE, L.E.A., JACOBS, A., JANSEN VAN RENSBURG, C., LAMANI, S., LOTZ, L.N., LOUW, S.VDM., LYLE, R., MALAN, A., MARAIS, M., NEETHLING, J.A., NXELE, T., PLISKO, D., PRENDINI, L., RINK, A.N., SWART, A., THERON, P., TRUTER, M., UECKERMANN, E., UYS, V.M., VILLET, M.H., WILLOWS-MUNROW, S. & WILSON, J.R.U. 2016. Soil biota in a megadiverse country: current knowledge and future research directions in South Africa. Pedobiologia 59: 129–174.


NEETHLING, J.A. & HADDAD, C.R. 2016. A systematic revision of the South African pseudoscorpions of the family Geogarypidae (Arachnida: Pseudoscorpiones). Indago 32: 1 – 80.


QUIÑONES-LEBRÓN, S.G., KRALJ-FIŠER, S., GREGORIC, M., LOKOVŠEK, T., CANDEK, K., HADDAD, C.R. & KUNTNER, M. 2016. Potential costs of heterospecific sexual interactions in golden orbweb spiders (Nephila spp.). Scientific Reports 6(#36908): 1–6.


SVOJANOVSKÁ, H., NGUYEN, P., HIRMAN, M., TUF, I.H., WAHAB, R.A., HADDAD, C.R. & ŠTÁHLAVSKÝ, F. 2016. Karyotype evolution in harvestmen of the suborder Cyphophthalmi (Opiliones). Cytogenetic and Genome Research 148: 227-236.




HADDAD, C.R. & MBO, Z. 2017. A new species of the endemic South African spider genus Austrachelas (Araneae: Gallieniellidae) and first description of the male of A. bergi. Zootaxa 4323: 119–124.

REZÁC, M., KREJCÍ, T., GOODACRE, S., HADDAD, C.R. & REZÁCOVÁ, V. 2017. Morphological and functional diversity of minor ampullate glands in spiders from the superfamily Amaurobioidea (Entelegynae: RTA clade). Journal of Arachnology 45: 198–208.



DIPPENAAR-SCHOEMAN, A.S., HADDAD, C.R., LYLE, R., LOTZ, L.N., FOORD, S.H., JOCQUÉ, R. & WEBB, P. 2018. South African National Survey of Arachnida: A checklist of the spiders (Arachnida, Araneae) of the Tswalu Kalahari Reserve in the Northern Cape province, South Africa. Koedoe 60(#a1486): 1–11.

HADDAD, C.R. & BUTLER, V.P. 2018. Ground-dwelling spider assemblages in contrasting habitats in the central South African Grassland Biome. Koedoe 60 (#a1482): 1–12.

LYLE, R. & HADDAD, C.R. 2018. Jocquestus gen. nov., a new genus of trachelid sac spiders from the Afrotropical Region (Arachnida: Araneae). Zootaxa 4471: 309–333.

PEKÁR, S., BOCÁNEK, O., MICHÁLEK, O., PETRÁKOVÁ, L., HADDAD, C.R., ŠEDO, O. & ZDRÁHAL, Z. (in press). Venom gland size and venom composition – essential trophic adaptations of venomous predators: the case study using spiders. Molecular Ecology.

ŠTÁHLAVSKÝ, F., JUST, P., OPATOVA, V., LOTZ, L.N. & HADDAD, C.R. 2018. Molecular technique reveals high variability of 18S rDNA distribution in harvestmen (Opiliones, Phalangiidae) from South Africa. Comparative Cytogenetics 12: 41–59.

WANG, C.X., LI, S.Q. & HADDAD, C.R. 2018. A new species of the spider genus Cangoderces (Araneae, Telemidae) from South Africa. African Invertebrates 59: 37–45.

Area(s) of Interest

African Arachnology. Various aspects of arachnid systematics and ecology are studied. The main focus is on the systematics and ecology of Afrotropical corinnid sac spiders, particularly ant-mimics in the subfamily Castianeirinae. This is one of the most poorly studied spider families in the region, and generic revisions presently underway indicate that the total diversity of this group is likely to increase at least five-fold. 

Other research areas include studies of spider biodiversity in habitats in the Free State Province, agroecosystems, and various conservancies. Biodiversity and ecological work focuses on the spider diversity of the Ndumo Game Reserve in Maputaland, KwaZulu-Natal, where more than 500 species of spiders have been recorded so far. Work on the role of spiders as predators of various insect pests in crops, and of termites in grassland, also receives attention. Collaborative research includes studies on the impacts of pesticides on spiders in crops, spider biology, particularly of termitophagous and myrmecophagous spiders, and karyology of South African arachnids.

Courses Presented

ENTO3714 - Advanced Insect Ecology (Sole lecturer). 

ZLGY/ENTO6814 - Scientific techniques (Part).

ENTO6844 – Arachnology capita selecta (Sole lecturer).



Elfrieda van den Berg (Marketing Manager)
T: +27 51 401 2531

Dilahlwane Mohono (Faculty Officer)
T: +27 58 718 5284

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